Joan Roughgarden is Professor Emerita of Biology at Stanford University and a Research Associate at the Hawaii Institute of Marine Biology at the University of Hawaii.

Joan Roughgarden is an evolutionary biologist, ecologist, and author whose work bridges biology, behavior, and social theory. She is Professor Emerita of Biology at Stanford University and a Research Associate at the Hawaii Institute of Marine Biology at the University of Hawaii. Over her career, she has contributed foundational research in evolutionary ecology, population genetics, and the evolution of social behavior. Roughgarden is widely known for her critiques of sexual selection theory and for advancing social selection as an alternative framework emphasizing cooperation and diversity in reproduction. Her books, including Evolution’s Rainbow and The Genial Gene, have shaped ongoing debates about sex, gender, and sexuality in both biology and society, positioning her as a leading voice in expanding evolutionary perspectives on diversity.

Events with Joan Roughgarden

Thursday November 13 at 12pm ET

[Live Podcast] Evolution’s Rainbow, Revisited: Sex, Gender, and Evolution

Free Live Discussion and Q&A with the Speakers

Groups with Joan Roughgarden

No groups with Joan Roughgarden

Authored by Joan Roughgarden

November 25, 2025

Evolution's Rainbow, Revisited: Sex, Gender, and Evolution

November 11, 2025

Sex, Gender Diversity, and Evolution: Introduction

To advance both science and society, we must reaffirm sex as a biological binary while expanding gender as an evolutionary spectrum of bodies and behaviors.