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Evolutionary Mismatch and How To Evaluate It: A Basic Tutorial

Evolutionary mismatch is a state of disequilibrium whereby an organism that evolved in one environment develops a phenotype that is harmful to its fitness or well-being in another environment.

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There is grandeur in this view of life, with its several powers, having been originally breathed into a few forms or into one; and that, whilst this planet has gone cycling on according to the fixed law of gravity, from so simple a beginning endless forms most beautiful and most wonderful have been, and are being, evolved.

- Charles Darwin, On the Origin of Species (1859)
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Evolutionary Science in Joyce’s Ulysses

James Joyce developed a writing technique that mirrored advances in the evolutionary science of his day and these insights are present in his novel. To explore this link, we can begin by looking at the most direct references to evolution science. Amidst the range of references to cultural figures in Ulysses, Charles Darwin makes a number of appearances, most notably in the fourteenth chapter, Oxen of the Sun.

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